Fungi Chapter 31 • Overview: Mighty Mushrooms • Fungi – Are diverse and widespread – Are essential for the well-being of most terrestrial ecosystems because.
Download ReportTranscript Fungi Chapter 31 • Overview: Mighty Mushrooms • Fungi – Are diverse and widespread – Are essential for the well-being of most terrestrial ecosystems because.
Fungi Chapter 31 • Overview: Mighty Mushrooms • Fungi – Are diverse and widespread – Are essential for the well-being of most terrestrial ecosystems because they break down organic material and recycle vital nutrients Figure 31.1 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Decomposers • Fungi are well adapted as decomposers of organic material – Performing essential recycling of chemical elements between the living and nonliving world Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Nutrition Despite their diversity – Fungi share some key traits • Fungi are heterotrophs – But do not ingest their food • Fungi secrete into their surroundings exoenzymes that break down complex molecules – And then absorb the remaining smaller compounds Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Fungi exhibit diverse lifestyles – Decomposers – Parasites – Mutualistic symbionts Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Fungi consist of – Mycelia, networks of branched hyphae adapted for absorption • Most fungi – Have cell walls made of chitin Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Body Structure • The morphology of multicellular fungi – Enhances their ability to absorb nutrients from their surroundings Reproductive structure. The mushroom produces tiny cells called spores. Hyphae. The mushroom and its subterranean mycelium are a continuous network of hyphae. Spore-producing structures 20 m Figure 31.2 Mycelium Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Some fungi – Have hyphae divided into cells by septa, with pores allowing cell-to-cell movement of materials • Coenocytic fungi – Lack septa Cell wall Cell wall Nuclei Pore Septum Figure 31.3a, b (a) Septate hypha Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Nuclei (b) Coenocytic hypha • Some unique fungi – Have specialized hyphae that allow them to penetrate the tissues of their host Nematode Hyphae 25 m (a) Hyphae adapted for trapping and killing prey Plant cell wall Fungal hypha (b) Haustoria Haustorium Figure 31.4a, b Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Plant cell Plant cell plasma membrane • Fungi produce spores through sexual or asexual life cycles • Fungi propagate themselves – By producing vast numbers of spores, either sexually or asexually Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • The generalized life cycle of fungi Key Heterokaryotic stage Haploid (n) Heterokaryotic (unfused nuclei from different parents) PLASMOGAMY (fusion of cytoplasm) Diploid (2n) KARYOGAMY (fusion of nuclei) Spore-producing structures Spores SEXUAL REPRODUCTION ASEXUAL REPRODUCTION Zygote Mycelium MEIOSIS GERMINATION GERMINATION Spore-producing structures Spores Figure 31.5 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Sexual Reproduction • The sexual life cycle involves – Cell fusion, plasmogamy – Nuclear fusion, karyogamy • An intervening heterokaryotic stage – Occurs between plasmogamy and karyogamy in which cells have haploid nuclei from two parents • The diploid phase following karyogamy – Is short-lived and undergoes meiosis, producing haploid spores Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Many fungi that can reproduce asexually – Grow as mold, sometimes on fruit, bread, and other foods 2.5 m Figure 31.6 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Other asexual fungi are yeasts – That inhabit moist environments – Which produce by simple cell division 10 m Parent cell Bud Figure 31.7 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Many molds and yeasts have no known sexual stage – Mycologists have traditionally called these deuteromycetes, or imperfect fungi Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Fungi descended from an aquatic, singlecelled, flagellated protist • Systematists now recognize Fungi and Animalia as sister kingdoms – Because fungi and animals are more closely related to each other than they are to plants or other eukaryotes Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings The Origin of Fungi • Molecular evidence – Supports the hypothesis that fungi and animals diverged from a common ancestor that was unicellular and bore flagella • Fungi probably evolved – Before the colonization of land by multicellular organisms Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • The oldest undisputed fossils of fungi – Are only about 460 million years old 50 m Figure 31.8 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Arbuscular mycorrhizal fungi Sac fungi Club fungi Ascomycota Basidiomycota Zygomycota Chytridiomycota Zygote fungi Chytrids Glomeromycota • The phylogeny of fungi Figure 31.9 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • A review of fungal phyla Table 31.1 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Symbionts • Fungi form symbiotic relationships with – Plants, algae, and animals • Mycorrhizae – Are mutually beneficial relationships between fungi and plant roots – 90% of all plant species have mycorrhizae relationships with fungi. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings The Move to Land • Fungi were among the earliest colonizers of land – Probably as symbionts with early land plants – The mycorrhizal symbiosis often increases the plant's uptake of inorganic compounds, such as nitrate and phosphate from soils – The fungal partners may also mediate plantto-plant transfer of carbohydrates and other nutrients. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Mycorrhizae • Mycorrhizae – Are enormously important in natural ecosystems and agriculture – Increase plant productivity EXPERIMENT Researchers grew soybean plants in soil treated with fungicide (poison that kills fungi) to prevent the formation of mycorrhizae in the experimental group. A control group was exposed to fungi that formed mycorrhizae in the soybean plants’ roots. RESULTS The soybean plant on the left is typical of the experimental group. Its RESULTS stunted growth is probably due to a phosphorus deficiency. The taller, healthier plant on the right is typical of the control group and has mycorrhizae. Figure 31.21 CONCLUSION These results indicate that the presence of mycorrhizae benefits a soybean plant and support the hypothesis that mycorrhizae enhance the plant’s ability to take up phosphate and other needed minerals. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Fungus-Animal Symbiosis • Some fungi share their digestive services with animals – Helping break down plant material in the guts of cows and other grazing mammals Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Many species of ants and termites – Take advantage of the digestive power of fungi by raising them in “farms” Figure 31.22 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Lichens • Lichens – Are a symbiotic association of millions of photosynthetic microorganisms held in a mass of fungal hyphae (a) A fruticose (shrub-like) lichen Figure 31.23a–c (b) A foliose (leaf-like) lichen Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings (c) Crustose (crust-like) lichens • The fungal component of a lichen – Is most often an ascomycete • Algae or cyanobacteria – Occupy an inner layer below the lichen surface Ascocarp of fungus Soredia Fungal hyphae Algal layer Algal cell Figure 31.24 10 m Fungal hyphae Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Pathogens • About 30% of known fungal species – Are parasites, mostly on or in plants Figure 31.25a–c (a) Corn smut on corn Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings (b) Tar spot fungus on maple leaves (c) Ergots on rye Practical Uses of Fungi • Some of the fungi that attack food crops – Are toxic to humans • Genetic research on fungi – Is leading to applications in biotechnology • Humans eat many fungi – And use others to make cheeses, alcoholic beverages, and bread Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Antibiotics produced by fungi – Treat bacterial infections Staphylococcus Penicillium Zone of inhibited growth Figure 31.26 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Chapter 33 Invertebrates • Overview: Life Without a Backbone • Invertebrates – Are animals that lack a backbone – Account for 95% of known animal species Figure 33.1 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings How Animals are Built • Nearly all animals follow a physical plan. There are three kinds. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Asymmetry The arrangement of body parts without central axis or point. • Asymmetrical organisms do not develop complex communication, sensory or motor functions Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Radial Symmetry • The arrangement of body parts such that any plane passing through the oralaboral axis divides the animal into mirror images • Allows for better specialization of sensory feeding and motor structure Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Bilateral Symmetry • The arrangement of body parts such that a single plane passing through the oral-aboral axis divides the animal into mirror images • Allows for advanced specialization of sensory, feeding and motor function which usually occurs in a distinct head Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Chordata Echinodermata Other bilaterians (including Nematoda, Arthropoda, Mollusca, and Annelida) Porifera Cnidaria • A review of animal phylogeny Deuterostomia Bilateria Eumetazoa Ancestral colonial choanoflagellate Figure 33.2 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Phylum Porifera • Sponges are sessile and have a porous body and choanocytes • Cellular – lack tissue sponges have no germ layers, no symmetry, no cephalization, no body cavity, and no segmentation • Reproduce sexually or asexually 5000 known species • Sponges, phylum Porifera – Live in both fresh and marine waters Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Sponges are suspension feeders – Capturing food particles suspended in the water that passes through their body 5 Choanocytes. The spongocoel is lined with feeding cells called choanocytes. By beating flagella, the choanocytes create a current that draws water in through the porocytes. Azure vase sponge (Callyspongia plicifera) 4 Spongocoel. Water passing through porocytes enters a cavity called the spongocoel. 3 Porocytes. Water enters the epidermis through channels formed by porocytes, doughnut-shaped cells that span the body wall. 2 Epidermis. The outer layer consists of tightly packed epidermal cells. Figure 33.4 Flagellum Collar Food particles in mucus Choanocyte Osculum Phagocytosis of food particles Spicules Water flow 1 Mesohyl. The wall of this simple sponge consists of two layers of cells separated by a gelatinous matrix, the mesohyl (“middle matter”). Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Amoebocyte 6 The movement of the choanocyte flagella also draws water through its collar of fingerlike projections. Food particles are trapped in the mucus coating the projections, engulfed by phagocytosis, and either digested or transferred to amoebocytes. 7 Amoebocyte. Amoebocytes transport nutrients to other cells of the sponge body and also produce materials for skeletal fibers (spicules). Phylum Cnidaria • Jellyfish, sea anemones, coral • Radial symmetry - polyps & medusa • Bodies contain “jelly” (mesoglea) • Specialized stinging cells Sea Anemone Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Tissue - lack organs have radial symmetry and 2 germ layers • lack cephalization, body cavity, & segmentation • Reproduce sexually or asexually 11,000 known species 4 groups - hydroids, box jellyfish, jellyfish, corals/anemones Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • The basic body plan of a cnidarian – Is a sac with a central digestive compartment, the gastrovascular cavity • A single opening – Functions as both mouth and anus Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • There are two variations on this body plan – The sessile polyp and the floating medusa Polyp Medusa Mouth/anus Tentacle Gastrovascular cavity Gastrodermis Mesoglea Body stalk Epidermis Tentacle Mouth/anus Figure 33.5 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Cnidarians are carnivores – That use tentacles to capture prey • The tentacles are armed with cnidocytes – Unique cells that function in defense and the capture of prey Prey Tentacle “Trigger” Discharge Of thread Nematocyst Figure 33.6 Coiled thread Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cnidocyte – Hydrozoa, Scyphozoa, Cubozoa, and Anthozoa – http://bcs.whfreeman.com/thelifewire/content/chp32/32020.html (a) These colonial polyps are members of class Hydrozoa. (b) Many species of jellies (class Scyphozoa), including the species pictured here, are bioluminescent. The largest scyphozoans have tentacles more than 100 m long dangling from a bell-shaped body up to 2 m in diameter. Figure 33.7a–d Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings (c) The sea wasp (Chironex fleckeri) is a member of class Cubozoa. Its poison, which can subdue fish and other large prey, is more potent than cobra venom. (d) Sea anemones and other members of class Anthozoa exist only as polyps. • Most animals have bilateral symmetry • The vast majority of animal species belong to the clade Bilateria – Which consists of animals with bilateral symmetry and triploblastic development Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Phylum Platyhelminthes • Free living flatworms, tapeworms, flukes • Well defined nervous, muscular, excretory, and reproductive systems • Many species have parasitic life style Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Tapeworm Platyhelminthes • Have Organ system • Platyhelminthes have bilateral symmetry and 3 germ layers • Cephalization • Lack body cavity and segmentation • Reproduce sexually and a sexually • Flat body allows for gas exchange, must occupy wet environments Examples - tapeworms, flukes (both endoparasites) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Turbellarian • Turbellarians – Are nearly all free-living and mostly marine Figure 33.9 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • The best-known turbellarians, commonly called planarians – Have light-sensitive eyespots and centralized nerve nets Pharynx. The mouth is at the tip of a muscular pharynx that extends from the animal’s ventral side. Digestive juices are spilled onto prey, and the pharynx sucks small pieces of food into the gastrovascular cavity, where digestion continues. Digestion is completed within the cells lining the gastrovascular cavity, which has three branches, each with fine subbranches that provide an extensive surface area. Undigested wastes are egested through the mouth. Gastrovascular cavity Eyespots Figure 33.10 Ganglia. Located at the anterior end of the worm, near the main sources of sensory input, is a pair of ganglia, dense clusters of nerve cells. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Ventral nerve cords. From the ganglia, a pair of ventral nerve cords runs the length of the body. • Trematodes that parasitize humans – Spend part of their lives in snail hosts 1 Mature flukes live in the blood vessels of the human intestine. A female fluke fits into a groove running the length of the larger male’s body, as shown in the light micrograph at right. Male Female 1 mm 5 These larvae penetrate the skin and blood vessels of humans working in irrigated fields contaminated with infected human feces. 2 Blood flukes reproduce sexually in the human host. The fertilized eggs exit the host in feces. 3 The eggs develop in water into ciliated larvae. These larvae infect snails, the intermediate hosts. Figure 33.11 4 Asexual reproduction within a snail results in another type of motile larva, which escapes from the snail host. Snail host Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Phylum Mollusca • Large phylum with over 50,000 species • Snails, limpets, slugs, sea hares, mussels, oysters, squid, octopus • Species often have a muscular foot, a mantle, and a shell Snail Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Mollusca • Organ System • Mollusca have bilateral symmetry and 3 germ layers • Cephalization • Coelom • No segmentation • Sexual (some hermaphroditic) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • All molluscs have a similar body plan with three main parts – A muscular foot – A visceral mass – A mantle Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Gastropods • About three-quarters of all living species of molluscs – Belong to class Gastropoda (a) A land snail Figure 33.18a, b (b) A sea slug. Nudibranchs, or sea slugs, lost their shell during their evolution. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cephalopods • Class Cephalopoda includes squids and octopuses – Carnivores with beak-like jaws surrounded by tentacles of their modified foot Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cephalopods • Most octopuses – Creep along the sea floor in search of prey – https://www.youtube.com/watch?v=AP_dpbTb ess&feature=related Figure 33.22a Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings (a) Octopuses are considered among the most intelligent invertebrates. • Squids use their siphon – To fire a jet of water, which allows them to swim very quickly Figure 33.22b (b) Squids are speedy carnivores with beaklike jaws and well-developed eyes. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Bivalves • Molluscs of class Bivalvia – Include many species of clams, oysters, mussels, and scallops – Have a shell divided into two halves Figure 33.20 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Phylum Annelida • Earthworms, featherdusters, leeches • Elongate cylindrical bodies in a series of segments • Well developed organ systems Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Leech Annelida • Organ System • Annelida have bilateral symmetry and 3 germ layers • Cephalization • Coelom and segmentation • Sexual reproduction (hermaphroditic) 16,500 known specices Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Annelida • Hydrostatic Skeleton • Circulatory system with 5 hearts • Excretory system including Nephridia • Nervous system has ganglionic brain and repeating ganglia • Digestive system is tubular gut with specialized parts Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Earthworms eat their way through the soil, extracting nutrients as the soil moves through the alimentary canal – Which helps till the earth, making earthworms valuable to farmers Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Leeches • Members of class Hirudinea – Are blood-sucking parasites, such as leeches – Used in medicinal purposes release Anesthetic and Anticoagulant at site of bite Figure Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings 33.25 Phylum Arthropoda • Hugely diverse and numerous phylum with over a million species • Have a tough exoskeleton • Jointed appendages and segmented body plan • Crabs, barnacles, insects, spiders, shrimp, centipedes Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Lobster • Organ System • Arthropoda have bilateral symmetry and 3 germ layers • Cephalization • Coelom • Segmentation • Sexual 1,000,000 known species Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Evolution of Arthropods • A hard exoskeleton • Presence of jointed appendages a complex nervous system with a brain connected to a ventral solid nerve cord • A unique respiratory system that emplys a variety of respiratory organs • A complex, yet adaptable,life cycle Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Early arthropods, such as trilobites – Showed little variation from segment to segment Figure 33.28 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings General Characteristics of Arthropods • The diversity and success of arthropods – Are largely related to their segmentation, hard exoskeleton, and jointed appendages Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • As arthropods evolved – The segments fused, and the appendages became more specialized • The appendages of some living arthropods – Are modified for many different functions Cephalothorax Abdomen Antennae (sensory reception) Head Thorax Swimming appendages Walking legs Figure 33.29 Pincer (defense) Mouthparts (feeding) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Millipedes, class Diplopoda – Have a large number of legs • Each trunk segment – Has two pairs of legs Figure 33.33 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Centipedes, class Chilopoda – Are carnivores with jaw-like mandibles – Have one pair of legs per trunk segment Figure 33.34 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cheliceriforms • Cheliceriforms, subphylum Cheliceriformes – Are named for clawlike feeding appendages called chelicerae – Include spiders, ticks, mites, scorpions, and horseshoe crabs Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Lack antennae • Chelicerata - distintive mouth parts • Many capture and sting insect prey • Others are ectoparisites • Horseshoe crabs feed on detritous • protosomes • Sexual reproduction only 70,000 species Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Most of the marine cheliceriforms are extinct – But some species survive today, including the horseshoe crabs Figure 33.30 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Most modern cheliceriforms are arachnids – A group that includes spiders, scorpions, ticks, and mites 50 µm (a) Scorpions have pedipalps that are pincers (b) Dust mites are ubiquitous scavengers in (c) Web-building spiders are generally specialized for defense and the capture of human dwellings but are harmless except most active during the daytime. food. The tip of the tail bears a poisonous to those people who are allergic to them stinger. (colorized SEM). Figure 33.31a–c Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Arachnids • Lack antennae • Cephalothorax and abdomen • Eight Walking legs • Carnivorous - liquid diet of bood or predigested prey • Simple eyes each with a single lens many spiders have eight eyes Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Arachnids have an abdomen and a cephalothorax – Which has six pairs of appendages, the most anterior of which are the chelicerae Intestine Digestive gland Stomach Heart Brain Eyes Poison gland Ovary Anus Book lung Spinnerets Figure 33.32 Silk gland Sperm Gonopore (exit for eggs) receptacle Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Chelicera Pedipalp Insects • Subphylum Hexapoda, insects and their relatives – Are more species-rich than all other forms of life combined – Live in almost every terrestrial habitat and in fresh water Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Many insects have some thoracic appendages modified for flight • Insects have a single pair of antenna, three pairs of legs, and in many cases, two wings Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • The internal anatomy of an insect – Includes several complex organ systems The insect body has three regions: head, thorax, and abdomen. The segmentation of the thorax and abdomen are obvious, but the segments that form the head are fused. Abdomen Thorax Head Compound eye Heart. The insect heart drives hemolymph through an open circulatory system. Cerebral ganglion. The two nerve cords meet in the head, where the ganglia of several anterior segments are fused into a cerebral ganglion (brain). The antennae, eyes, and other sense organs are concentrated on the head. Antennae Ovary Figure 33.35 Malpighian tubules. Anus Metabolic wastes are removed from the Vagina hemolymph by excretory organs called Malpighian tubules, which are outpocketings of the digestive tract. Tracheal tubes. Gas exchange in insects is accomplished by a tracheal system of branched, chitin-lined tubes that infiltrate the body and carry oxygen directly to cells. The tracheal system opens to the outside of the body through spiracles, pores that can control air flow and water loss by opening or closing. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Nerve cords. The insect nervous system consists of a pair of ventral nerve cords with several segmental ganglia. Dorsal artery Crop Insect mouthparts are formed from several pairs of modified appendages. The mouthparts include mandibles, which grasshoppers use for chewing. In other insects, mouthparts are specialized for lapping, piercing, or sucking. • Flight is obviously one key to the great success of insects • An animal that can fly – Can escape predators, find food, and disperse to new habitats much faster than organisms that can only crawl Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Many insects – Undergo metamorphosis during their development • In incomplete metamorphosis, the young, called nymphs – Resemble adults but are smaller and go through a series of molts until they reach full size Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Insects with complete metamorphosis – Have larval stages specialized for eating and growing that are known by such names as maggot, grub, or caterpillar • The larval stage – Looks entirely different from the adult stage Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Metamorphosis from the larval stage to the adult stage – Occurs during a pupal stage (a) Larva (caterpillar) (b) Pupa (c) Pupa (d) Emerging adult Figure 33.6a–e Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings (e) Adult Ecological Role of Insects • Pollination • decomposition of organic material • recycling of carbon, nitrogen, and other essential nutrients • Control of population of harmful invertebrate species • Direct production of certain foods manufacture useful products such as silk Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Phylum Echinoderm • Entirely marine group containing sea stars, brittle stars, urchins, sand dollars, sea cucumbers, sea lilies • Marked radial symmetry • Unique water vascular system - tube feet Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Sea Urchin Echinoderms • Organ systems • Echinoderms have bilateral symmetry as larvae and radial symmetry as adults • 3 germ layers • No cephalization or Segmentation • Coelum • Sexual (some hermaphroditic; asexual by regeneration) 6500 known species Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Echinoderms and chordates are deuterostomes • At first glance, sea stars and other echinoderms, phylum Echinodermata – May seem to have little in common with phylum Chordata, which includes the vertebrates Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Chordates and echinoderms share characteristics of deuterostomes – Radial cleavage – Development of the coelom from the archenteron – Formation of the mouth at the end of the embryo opposite the blastopore Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Unique to echinoderms is a water vascular system – A network of hydraulic canals branching into tube feet that function in locomotion, feeding, and gas exchange A short digestive tract runs from the mouth on the bottom of the central disk to the anus on top of the disk. Central disk. The central disk has a nerve ring and nerve cords radiating from the ring into the arms. Digestive glands secrete digestive juices and aid in the absorption and storage of nutrients. Figure 33.39 Radial canal. The water vascular system consists of a ring canal in the central disk and five radial canals, each running in a groove down the entire length of an arm. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Stomach Anus Spine Gills Ring canal Gonads Radial nerve The surface of a sea star is covered by spines that help defend against predators, as well as by small gills that provide gas exchange. Madreporite. Water can flow in or out of the water vascular system into the surrounding water through the madreporite. Ampulla Podium Tube feet Branching from each radial canal are hundreds of hollow, muscular tube feet filled with fluid. Each tube foot consists of a bulb-like ampulla and suckered podium (foot portion). When the ampulla squeezes, it forces water into the podium and makes it expand. The podium then contacts the substrate. When the muscles in the wall of the podium contract, they force water back into the ampulla, making the podium shorten and bend. Sea Stars • Sea stars, class Asteroidea – Have multiple arms radiating from a central disk • The undersurfaces of the arms – Bear tube feet, each of which can act like a suction disk Figure 33.40a (a) A sea star (class Asteroidea) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Brittle Stars • Brittle stars have a distinct central disk – And long, flexible arms Figure 33.40b (b) A brittle star (class Ophiuroidea) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Sea Cucumbers • Feather stars – Crawl about using their long, flexible arms Figure 33.40d (d) A feather star (class Crinoidea) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Sea Daisies • Sea daisies were discovered in 1986 – And only two species are known Figure 33.40f (f) A sea daisy (class Concentricycloidea) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Phylum Chordata • Includes animals with a “backbone” • 7 Classes within the Subphylum Vertebrata Sharks, bony fish, lamprey, amphibians, reptiles, birds, mammals • We will talk about them in detail next time………. • http://quizlet.com/2981733/biologyflash-cards/ Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings